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In human endothelial cells, beetroot methanolic extract suppressed NF-κB activation and reduced expression of TNF-α, toll-like receptor-4 (TLR-4), and IL-1β, while upregulating nuclear factor erythroid 2–related factor 2 (Nrf2), endothelial nitric oxide synthase (eNOS), glutathione peroxidase (GPX), and glycogen synthase kinase-3β (GSK-3β), indicating coordinated anti-inflammatory and cytoprotective signaling (Al-Harbi et al., 2021b)